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Paul Bettany as Charles Darwin |
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Jennifer Connelly as Emma Darwin |
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Martha West as Anne Darwin |
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Guy Henry as Technician |
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Jeremy Northam as John Brodie-Innes |
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Toby Jones as Thomas Henry Huxley |
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Benedict Cumberbatch as Joseph Dalton Hooker |
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Jim Carter as Parslow |
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Robert Glenister as Dr. Holland |
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Ian Mercer as Goodman |
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Bill Paterson as Dr. Gully |
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Teresa Churcher as Mrs. Davies |
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Richard Ridings as Thatcher |
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Freya Parks as Etty Darwin |
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Ian Kelly as Captain Fitzroy |
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Anabolena Rodriguez as Fuegia Basket |
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Paul Campbell as Boat Memory |
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Zak Davies as Jemmy Button |
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Christopher Dunkin as George Darwin |
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Gene Goodman as Franky Darwin |
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Harrison Sansostri as Lenny Darwin |
| Production | Jamie Laurenson | Executive Producer |
| Production | Jeremy Thomas | Producer |
| Directing | Jon Amiel | Director |
| Writing | John Collee | Screenplay |
| Writing | John Collee | Screenstory |
| Production | Pip Ayers | Producer |
| Writing | Randal Keynes | Novel |
| Camera | Jess Hall | Director of Photography |
| Editing | Melanie Oliver | Editor |
| Art | Laurence Dorman | Production Design |
| Sound | Christopher Young | Original Music Composer |
| Production | David M. Thompson | Executive Producer |
| Production | Christina Yao | Executive Producer |
| Production | Celestia Fox | Casting |
| Writing | Jon Amiel | Screenstory |
| Production | Nick O'Hagan | Producer |
| Production | Janice Eymann | Executive Producer |
| Production | Peter Watson | Executive Producer |
| Production | Tom Waller | Line Producer |
| Costume & Make-Up | Veronica McAleer | Makeup Artist |
| Sound | Paul Hanks | Foley Artist |
| Sound | Chris Burdon | Sound Re-Recording Mixer |
| Sound | Ruth Sullivan | Foley Artist |
| Costume & Make-Up | Ashley Johnson | Makeup Artist |
| Sound | Sue Harding | Foley Artist |
| Costume & Make-Up | Francesco Alberico | Hairstylist |
| Sound | Peter Gleaves | ADR Mixer |
| Sound | Jack Stew | Foley Artist |
| Costume & Make-Up | Vitchu Chavasit | Makeup Artist |
| Production | Sara Desmond | Unit Production Manager |
| Art | Gary Jopling | Art Direction |
| Art | Dominic Capon | Set Decoration |
| Costume & Make-Up | Louise Stjernsward | Costume Design |
| Art | Bill Crutcher | Supervising Art Director |
| Lighting | Lee Walters | Gaffer |
